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Pancheloniidae

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Pancheloniids Temporal range: Late Cretaceous-Holocene, 73–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N

Ctenochelys stenoporus skeletons
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Clade:	Panchelonioidea
Clade:	Pancheloniidae
Subgroups
See text.

Pancheloniidae is a clade of sea turtles^[1] It is defined as all turtles more closely related to cheloniid sea turtles than to dermochelyid ("leatherback") sea turtles.^[2]

Genera

The following genera are placed here:

Historic taxon placement

The following list of non-cheloniid pancheloniid species was published by Hirayama and Tong in 2003^[4]

†Allopleuron hoffmani,^[4] later assigned to Cheloniidae
†Catapleura arcansaw^[4]
†Catapleura repanda^[4]
†Ctenochelys stenoporus^[4]
†Erquelinnesia gosseleti^[4]
†Glyptochelone suyckerbuykii^[4]
†Lytoloma cantabrigiens^[4]
†Osteopygis emarginatus^[4]
†Pampaemys meridionalis^[4]
†Peritresius ornatus^[4]
†Porthochelys laticeps^[4]
†Prionochelys nauta^[4]
†Retechelys platyops^[4]
†Tasbacka aldabergeni,^[4] later assigned to Cheloniidae
†Thinochelys lapisossea^[4]
†Toxochelys latiremis,^[4] later assigned to clade Durocryptodira.^[8]

Cladogram

Below is a cladogram showing the phylogenetic relationships of living and extinct sea turtles in the clade Pancheloniidae based on Lynch and Parham (2003)^[9] and Parham and Pyenson (2010).^[10]

Lytoloma fossil

Pancheloniidae

†Toxochelys

†Mexichelys

†Lophochleyinae

†Argillochelys

	†Erquelinnesla

	†Pacifichelys

	†Puppigerus

	Cheloniidae

References

^ Kear BP, Lee MS (March 2006). "A primitive protostegid from Australia and early sea turtle evolution". Biol. Lett. 2 (1): 116–9. doi:10.1098/rsbl.2005.0406. PMC 1617175. PMID 17148342.
^ Joyce W.G, Parham J.F, Gauthier J.A. Developing a protocol for the conversion of rank-based taxon names to phylogenetically defined clade names, as exemplified by turtles. J. Paleont. 2004;78:989–1013.
^ E.A. Zvonok; I.G. Danilov (2017). "A revision of fossil turtles from the Kiev clays (Ukraine, middle Eocene) with comments on the history of the collection of fossil vertebrates of A.S. Rogovich" (PDF). Proceedings of the Zoological Institute of the Russian Academy of Sciences. 321 (4): 485–516.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad Hirayama, Ren; Tong, Haiyan (2003). "Osteopygis (Testudines: Cheloniidae) from the Lower Tertiary of the Ouled Abdoun phosphate basin, Morocco". Palaeontology. 46 (5): 845–856. Bibcode:2003Palgy..46..845H. doi:10.1111/1475-4983.00322.^{[permanent dead link‍]}
^ Andrew D. Gentry (2018). "Prionochelys matutina Zangerl, 1953 (Testudines: Pan-Cheloniidae) from the Late Cretaceous of the United States and the evolution of epithecal ossifications in marine turtles". PeerJ. 6: e5876. doi:10.7717/peerj.5876. PMC 6215699. PMID 30402356.
^ Danilov, I.G.; Averianov, A.O.; Yarkov, A.A. (2010). "Itilochelys rasstrigin gen. et sp. nov., a new hard-shelled sea turtle (Cheloniidae sensu lato) from the Lower Palaeocene of Volgograd Province, Russia" (PDF). Proceedings of the Zoological Institute RAS. 314 (1): 24–41. doi:10.31610/trudyzin/2010.314.1.24.
^ "Pancheloniidae". Fossilworks. Retrieved 17 December 2021.
^ "Toxochelys". FossilWorks. Retrieved 17 December 2021.
^ Lynch, S.C.; Parham, J.F. (2003). "The first report of hard-shelled sea turtles (Cheloniidae sensu lato) from the Miocene of California, including a new species (Euclastes hutchisoni) with unusually plesiomorphic characters" (PDF). PaleoBios. 23 (3): 21–35.^{[permanent dead link‍]}
^ James F. Parham; Nicholas D. Pyenson (2010). "New Sea Turtle from the Miocene of Peru and the Iterative Evolution of Feeding Ecomorphologies since the Cretaceous". Journal of Paleontology. 84 (2): 231–247. Bibcode:2010JPal...84..231P. doi:10.1666/09-077R.1. S2CID 62811400.

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

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