Soldier beetle

Soldier beetle Temporal range: Albian–Present PreꞒ Ꞓ O S D C P T J K Pg N
Chauliognathus lugubris
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Elateriformia
Superfamily:	Elateroidea
Family:	Cantharidae Imhoff, 1856
Synonyms
Chauliognathidae

The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the red coats of early British soldiers, hence the common name. They are also known commonly as leatherwings because of their soft elytra.^[1]

Historically, these beetles were placed in a superfamily "Cantharoidea", which has been subsumed by the superfamily Elateroidea; the name is still sometimes used as a rankless grouping, including the families Cantharidae, Lampyridae, Lycidae, Omethidae (which includes Telegeusidae), Phengodidae, and Rhagophthalmidae.

Soldier beetles often feed on nectar and pollen as well as predating other small insects. The larvae are caterpillar like, dark colored, active and covered in fine hairs, earning them the name velvet worms. They feed on the ground and in foliage hunting eggs, small insects, snails and other small creatures. ^{[2] [3]}

The oldest described member of the family is Molliberus from the Early Cretaceous (early Albian) aged El Soplao amber from Cantabria, Spain, belonging to the tribe Cantharini in the subfamily Cantharinae. Other described genera include 6 from the early Late Cretaceous (early Cenomanian) aged Burmese amber, with 5 belonging to Cantharinae and one to Malthininae, and Katyacantharis, from the Cenomanian aged Agdzhakend amber of Azerbaijan, suggested to belong to Cantharinae. Indeterminate specimens have been reported from the Aptian aged Koonwarra fossil bed of the Strzelecki Group, Australia and the Barremian aged Lebanese amber.^[4][5]

Five subfamilies are normally accepted:

• tribe Cantharini
  • Cantharis
  • Rhagonycha
• tribe Podabrini
  • Podabrus

• tribe Chauliognathini
  • Belotus
  • Chauliognathus
• tribe Ichthyurini
  • Trypherus

• Afronycha Wittmer, 1949 - central-southern Africa
• Asilis Broun, 1893 - New Zealand
• Compsonycha
• Dysmorphocerus Solier, 1849
• Flabelloontelus
• Geigyella Wittmer, 1972 - New Guinea
• Hansasilis
• Heteromastix Boheman, 1858 - Australia
• Hyponotum
• Micronotum
• Neoontelus Wittmer, 1972 - New Zealand
• Oontelus Solier, 1849 - S. America
• Plectocephalon
• Plectonotum Gorham, 1891 - Americas

• tribe Malchinini
  • Macrocerus Motschulsky, 1845 - Europe (synonym Malchinus)
• tribe Malthinini
  • Caccodes Sharp, 1885 - Central America, Pacific islands
  • Malthinellus Kiesenwetter, 1874 - Japan
  • Malthinus Latreille, 1805 - Japan, Europe, N. America
• tribe Malthodini
  • Frostia Bert. ex Guill.
  • Malthodes Kiesenwetter, 1852 - mostly Europe, N. America & New Zealand
  • †Archaeomalthodes Hsiao et al. 2016 Burmese amber, Myanmar, Cenomanian

• tribe Silini
  • Cordylocera Guérin-Méneville, 1830
  • Silis Charpentier, 1825
• tribe Tytthonyxini
  • Tytthonyx LeConte, 1851

Large males of the soldier beetle exercise choice for larger females. Body size correlates with the abilities of males to secure females, and of females to evade males.^[6]

• List of Cantharidae genera