Lycium

Lycium
African boxthorn (Lycium ferocissimum)
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Solanales
Family:	Solanaceae
Tribe:	Lycieae
Genus:	Lycium L. (1753)[1]
Species[2]
101; see text
Synonyms[2]
Ascleia Raf. (1838) Cantalea Raf. (1838) Evoista Raf. (1838) Grabowskia Schltdl. (1832) Jasminoides Duhamel (1755) Oplukion Raf. (1838) Panzeria J.F.Gmel. (1791) Phrodus Miers (1849) Pukanthus Raf. (1838) Rhopalostigma Phil. (1860) Teremis Raf. (1838) Triliena Raf. (1838)

Lycium is a genus of flowering plants in the nightshade family, Solanaceae.^[3] The genus has a disjunct distribution around the globe, with species occurring on most continents in temperate and subtropical regions. South America has the most species, followed by North America and southern Africa. There are several scattered across Europe and Asia, and one is native to Australia.^[4] Common English names for plants of this genus include box-thorn,^[5] wolfberry, and desert-thorn.^[6] Plants of the World Online currently accepts 101 species.^[2] Other estimates are of 70^[4] to 80^[7][8] species.

The generic name Lycium is derived from the Greek word λυκιον (lykion), which was applied by Pliny the Elder (23-79) and Pedanius Dioscorides (ca. 40–90) to a plant known as dyer's buckthorn. It was probably a Rhamnus species and was named for Lycia (Λυκία), the ancient southern Anatolian region in which it grew.^[9][10] The berry is called lycii fructus ("lycium fruit") in old Latin pharmacological texts.

Lycium are shrubs, often thorny, growing 1 to 4 meters tall. The leaves are small, narrow, and fleshy, and are alternately arranged, sometimes in fascicles. Flowers are solitary or borne in clusters. The funnel-shaped or bell-shaped corolla is white, green, or purple in color. The fruit is a two-chambered, usually fleshy and juicy berry which can be red, orange, yellow, or black. It may have few seeds or many.^[5][7] Most Lycium have fleshy, red berries with over 10 seeds, but a few American taxa have hard fruits with two seeds.^[8]

While most Lycium are monoecious, producing bisexual flowers with functional male and female parts, some species are gynodioecious, with some individuals bearing bisexual flowers and some producing functionally female flowers.^[11]

Lycium has been known to European herbalists since ancient times, and species were traded from the Far East to Europe by the Romans, for example via Ariaca and the port of Barbarikon near today's Karachi, as mentioned in the Periplus of the Erythraean Sea. In his Naturalis historia, Pliny the Elder describes boxthorn as a medicinal plant, as does Pedanius Dioscorides in his P. Dioscoridae pharmacorum simplicium reique medicae.^[12]

In his 1753 publication Species Plantarum, Linnaeus describes three Lycium species: L. afrum, L. barbarum, and L. europaeum.^[12]

Lycium, particularly L. barbarum, have long been used in traditional Chinese medicine.^[13][14] The leaves and roots of other species of Lycium, such as L. europaeum, when mixed with water, have been used in folk medicine.^[15] The fruit of L. barbatum and L. chinense, known as goji berry, is commonly consumed as a dried fruit.^[13] The Chinese tonic gou qi zi ("wolfberry fruit") is made of the fruit of any of several Lycium species, and is used as a dietary supplement.^[13]

Lycium species mostly occur in arid and semi-arid climates, and a few are known from coastal zones in somewhat saline habitat types.^[4]

Invasive species include L. ferocissimum, which was introduced to Australia and New Zealand and has become a dense, thorny pest plant there. It injures livestock, harbors pest mammals and insects, and displaces native species.^[16]

101 species are accepted.^[2][17][18]

• Buchozia japonica (Thunb.) Callm. (as L. foetidum L.f. or L. japonicum Thunb.)^[17]

Wikimedia Commons has media related to Lycium.

Wikispecies has information related to Lycium.

• Project Lycieae Archived 2016-03-04 at the Wayback Machine, Amherst College