Heteractinida

Heteractinida; Temporal range: Cambrian–Permian PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Porifera
Informal group:	†"Heteractinida"; Hinde, 1887
Included groups
	See text
Synonyms
	Octactinellidae Hinde, 1887; Heteractinellidae Hinde, 1887; Wewokellida Croneis & Toomey, 1965;

Heteractinida is an extinct grade of Paleozoic (Cambrian–Permian) sponges, sometimes used as a class or order. They are most commonly considered paraphyletic with respect to Calcarea (calcareous sponges),^[1]^[2]^[3] though some studies instead argue that they are paraphyletic relative to Hexactinellida (glass sponges).^[4] Heteractinids can be distinguished by their six-pronged (snowflake-shaped) spicules, whose symmetry historically suggested a relationship with the triradial calcarean sponges.^[4]

The sponge-like Cambrian Hetairacyathidae may be related to heteractinids,^[5] though most instead consider hetairacyathids to be closer to archaeocyaths and radiocyaths.^[6] An example of a species in the Heteractinida class is the Gondekia Lancifer; the Gondekia lancifer specimen is only the second articulated specimen known and is more complete than the holotype. It has an intact skeleton of felted sexiradiates of three orders of spicules.^[7]

Subgroups

From Treatise on Invertebrate Paleontology:^[5]

Order †Octactinellida Hinde, 1887
- Family †Astraeospongiidae Miller, 1889 [Cambrian–Devonian]
- Family †Eiffeliidae Rigby, 1986 [Cambrian–Mississippian]
- Family †Nuchidae Pickett, 2002 [Cambrian]
- Family †Wewokellidae King, 1943 [Mississippian–Permian]

References

^ Pickett, J. (2002). "†Order Heteractinida Hinde, 1887". In Hooper, J.N.A.; Van Soest, R.W.M.; Willenz, P. (eds.). Systema Porifera: A Guide to the Classification of Sponges. Boston, MA: Springer. pp. 1121–1140. doi:10.1007/978-1-4615-0747-5_117. ISBN 978-0-306-47260-2.
^ Jain, Sreepat (2017). "Sponges". Fundamentals of Invertebrate Palaeontology. Springer Geology. New Delhi: Springer India. pp. 7–30. doi:10.1007/978-81-322-3658-0. ISBN 978-81-322-3656-6.
^ Nadhira, Ardianty; Sutton, Mark D.; Botting, Joseph P.; Muir, Lucy A.; Gueriau, Pierre; King, Andrew; Briggs, Derek E. G.; Siveter, David J.; Siveter, Derek J. (2019). "Three-dimensionally preserved soft tissues and calcareous hexactins in a Silurian sponge: implications for early sponge evolution". Royal Society Open Science. 6 (7): 190911. doi:10.1098/rsos.190911. ISSN 2054-5703. PMC 6689616. PMID 31417767.
^ ^a ^b Botting, J. P.; Butterfield, N. J.; Valentine, J. W. (2005). "Reconstructing early sponge relationships by using the Burgess Shale fossil Eiffelia globosa, Walcott". Proceedings of the National Academy of Sciences of the United States of America. 102 (5): 1554–1559. doi:10.1073/pnas.0405867102. JSTOR 3374469. PMC 547825. PMID 15665105.
^ ^a ^b Treatise on Invertebrate Paleontology Part E, Revised. Porifera, Volume 3: Classes Demospongea, Hexactinellida, Heteractinida & Calcarea, xxxi + 872 p., 506 fig., 1 table, 2004, available here. ISBN 0-8137-3131-3.
^ Treatise on Invertebrate Paleontology Part E, Revised. Porifera, Volumes 4 & 5: Hypercalcified Porifera, Paleozoic Stromatoporoidea & Archaeocyatha, liii + 1223 p., 665 figs., 2015, available here. ISBN 978-0-9903621-2-8.
^ Rigby, J. Keith (1994). "Well-Preserved Specimens of the Sponges Gondekia (Heteractinida) and Pseudohydnoceras (Hexactinellida), Middle Devonian of New York State". Journal of Paleontology. 68 (4): 727–734. ISSN 0022-3360.

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[1] Pickett, J. (2002). "†Order Heteractinida Hinde, 1887". In Hooper, J.N.A.; Van Soest, R.W.M.; Willenz, P. (eds.). Systema Porifera: A Guide to the Classification of Sponges. Boston, MA: Springer. pp. 1121–1140. doi:10.1007/978-1-4615-0747-5_117. ISBN 978-0-306-47260-2.

[2] Jain, Sreepat (2017). "Sponges". Fundamentals of Invertebrate Palaeontology. Springer Geology. New Delhi: Springer India. pp. 7–30. doi:10.1007/978-81-322-3658-0. ISBN 978-81-322-3656-6.

[3] Nadhira, Ardianty; Sutton, Mark D.; Botting, Joseph P.; Muir, Lucy A.; Gueriau, Pierre; King, Andrew; Briggs, Derek E. G.; Siveter, David J.; Siveter, Derek J. (2019). "Three-dimensionally preserved soft tissues and calcareous hexactins in a Silurian sponge: implications for early sponge evolution". Royal Society Open Science. 6 (7): 190911. doi:10.1098/rsos.190911. ISSN 2054-5703. PMC 6689616. PMID 31417767.

[ref_-4] Botting, J. P.; Butterfield, N. J.; Valentine, J. W. (2005). "Reconstructing early sponge relationships by using the Burgess Shale fossil Eiffelia globosa, Walcott". Proceedings of the National Academy of Sciences of the United States of America. 102 (5): 1554–1559. doi:10.1073/pnas.0405867102. JSTOR 3374469. PMC 547825. PMID 15665105.

[:0-5] Treatise on Invertebrate Paleontology Part E, Revised. Porifera, Volume 3: Classes Demospongea, Hexactinellida, Heteractinida & Calcarea, xxxi + 872 p., 506 fig., 1 table, 2004, available here. ISBN 0-8137-3131-3.

[:1-6] Treatise on Invertebrate Paleontology Part E, Revised. Porifera, Volumes 4 & 5: Hypercalcified Porifera, Paleozoic Stromatoporoidea & Archaeocyatha, liii + 1223 p., 665 figs., 2015, available here. ISBN 978-0-9903621-2-8.

[7] Rigby, J. Keith (1994). "Well-Preserved Specimens of the Sponges Gondekia (Heteractinida) and Pseudohydnoceras (Hexactinellida), Middle Devonian of New York State". Journal of Paleontology. 68 (4): 727–734. ISSN 0022-3360.

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