Carychium

Carychium
	Shell of Carychium minimum
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Ellobiida
Family:	Ellobiidae
Subfamily:	Carychiinae
Genus:	Carychium; O. F. Müller, 1773
Synonyms
	Auricella Jurine, 1817; Auriculina Moquin-Tandon, 1856; Carychium (Carychium) O.F. Müller, 1773· accepted, alternate representation; Carychium (Saraphia) Risso, 1826 (junior synonym); Saraphia Risso, 1826;

Carychium is a genus of very small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Ellobiidae.^[2]

Species

species within the genus Carychium include:

Carychium achimszulci Stworzewicz, 1999,^[3] Miocene
Carychium antiquum Braun, 1843, Miocence ^[4]
†Carychium apathyi Gaál, 1911,^[4] Miocene
Carychium arboreum Dourson, 2012^[5]
Carychium belizeense Jochum & Weigand, 2017 ^[6]
†Carychium berellense Laubrière & Carez, 1880,^[4] Paleocene
†Carychium bermudense Gulick, 1904,^[4] Pliocene
†Carychium bigeminatum (Deshayes, 1863),^[4] Paleocene
Carychium biondii Paulucci, 1882
Carychium boysianum Benson, 1864 ^[7]
Carychium carinatum Haufen, 1858 ^[8]
†Carychium cholnokyi Gaál, 1911,^[4] Miocene
Carychium clappi Hubricht, 1959
Carychium costaricanum Von Martens, 1898
†Carychium cylindroides Staadt, 1913,^[4] Paleocene
Carychium cymatoplax Pilsbry, 1901
†Carychium dhorni (Deshayes, 1863),^[4] Paleocene
†Carychium euboicum Schütt, 1988,^[3] Miocene
†Carychium eumicrum Bourguignat, 1857,^[3] Miocene
Carychium exiguum (Say, 1822)
Carychium exile H. C. Lea, 1842
†Carychium fischeri Boettger, 1903,^[4] Oligocene
Carychium floridanum Clapp, 1918
Carychium gibbum (Sandberger, 1875)^[9]
†Carychium gracile Sandberger, 1875,^[3] Miocene
Carychium hachijoensis Pilsbry, 1902
Carychium hardiei Jochum & Weigand, 2017 ^[6]
Carychium hellenicum Bank & Gittenberger, 1985
†Carychium hypermeces Cossmann, 1889,^[4] Paleocene
Carychium ibazoricum Bank & Gittenberger, 1985
Carychium indicum Benson, 1849 ^[10]
†Carychium interferens (Deshayes, 1863),^[4] Paleocene
Carychium jardineanum (Chitty, 1853) ^[11]
Carychium javanum von Möllendorff, 1897 ^[12]
Carychium lederi O. Boettger, 1880
Carychium loheri von Möllendorff, 1898 ^[13]
†Carychium majus Boettger, 1870,^[4] Miocene
Carychium mariae Paulucci, 1878
Carychium mexicanum Pilsbry, 1891
?†Carychium michaudi (Boissy, 1848),^[4] Paleocene
†Carychium michelini (Boissy, 1846),^[4] Paleocene
Carychium minimum ( O. F. Müller, 1774)^[14]
Carychium minusculum Gredler, 1887
†Carychium moenanum Wenz, 1917,^[4] Oligocene
Carychium nannodes G. H. Clapp, 1905
†Carychium nincki Cossmann, 1913,^[4] Eocene
Carychium nipponense Pilsbry & Hirase, 1904
Carychium noduliferum Reinhardt, 1877
†Carychium nouleti Bourguignat, 1857,^[4] Miocene/Pliocene
Carychium occidentale Pilsbry, 1891
†Carychium pachychilus Sandberger, 1875,^[4] Miocene
Carychium paganettii Zimmermann, 1925
Carychium panamaense Jochum, 2018 ^[15]
Carychium pessimum Pilsbry, 1902
†Carychium pseudotetrodon Strauch, 1977,^[3] Miocene
†Carychium puisseguri Truc, 1972,^[16] Pliocene
Carychium pulchellum Freyer 1855,^[17]
†Carychium quadridens (Andreae, 1884),^[4] Eocene
†Carychium remiensis (Boissy, 1848),^[4] Paleocene
†Carychium rhenanum Strauch, 1977,^[3] Pliocene
Carychium riparium Hubricht, 1978
†Carychium sandbergeri Handmann, 1887,^[4] Miocene/Pliocene
†Carychium schlickumi Strauch, 1977,^[18] Miocene/Pliocene
†Carychium schwageri Reuss, 1868,^[3] Miocene
?Carychium sianicum Caziot, 1910, subrecent
Carychium sibiricum Gerstfeldt, 1859
†Carychium sparnacense Deshayes, 1863,^[4] Paleocene
†Carychium starobogatovi Steklov, 1966,^[18] Miocene/Pliocene
Carychium stygium Call, 1897
†Carychium suevicum Boettger, 1877,^[3] Miocene
†Carychium surai Stworzewicz, 1999,^[3] Miocene
†Carychium tetrodon (Paladilhe, 1873),^[4] Paleocene
Carychium thailandicum Burch & Panha, 1998
Carychium tianmushanense Chen, 1992
Carychium tridentatum (Risso, 1826)
†Carychium vindobonense (Handmann, 1882),^[4] Pliocene
Carychium zarzaae Jochum & Weigand, 2017 ^[6]

References

Literature

R. A. Bank, E. Gittenberger: Notes on Azorean and European Carychium species (Gastropoda Basommatophora: Ellobiidae). In: Basteria. 49, Leiden 1985, S. 85–100.
A. Jochum: Evolution and diversity of the troglobitic Carychiidae – A morphological and phylogenetic investigation of the terrestrial ellobiiod genera, Carychium and Zospeum. In: The Malacologist. 57, London 2011, S. 16–18. (PDF)
F. Strauch: Die Entwicklung der europäischen Vertreter der Gattung Carychium O. F. Müller seit dem Miozän. In: Archiv für Molluskenkunde. 107 (4/6), Frankfurt am Main 1977, S. 149–193.
A. M. Weigand, A. Jochum, R. Slapnik, J. Schnitzler, E. Zarza, A. Klussmann-Kolb: Evolution of microgastropods (Ellobioidea, Carychiidae): integrating taxonomic, phylogenetic and evolutionary hypotheses. In: BMC Evolutionary Biology. 13, 2013, S. 18. (biomedcentral.com, PDF; 2,7 MB)
A. M. Weigand, M.-C. Goetze, A Jochum: Outdated but established?! Conchologically driven species delineations in microgastropods (Carychiidae, Carychium). In: Organisms Diversity and Evolution. 12, 2012, S. 377–386.
J. C. Nekola, M. Barthel: Morphometric analysis of the genus Carychium in the Great Lakes region. In: Journal of Conchology. 37, 2002, S. 515–531.
K. Bulman: Shell variability in Carychium tridentatum (Risso, 1826) and its importance for infraspecific taxonomy (Gastropoda, Pulmonata, Ellobiidae). In: Malakologische Abhandlungen des Museums für Tierkunde Dresden. 15, 1990, S. 37–50.
A. M. Weigand, A. Jochum, R. Slapnik, J. Schnitzler, E. Zarza, A. Klussmann-Kolb: Evolution of microgastropods (Ellobioidea, Carychiidae): integrating taxonomic, phylogenetic and evolutionary hypotheses. In: BMC Evolutionary Biology. 13, 2013, S. 18 (biomedcentral.com PDF; 2,7 MB)
O. F. Müller: Vermivm terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum et testaceorum, non marinorum, succincta historia. In: Voluminis Imi pars Ima. 1773, S. [1-33], 1–135. Havniæ & Lipsiæ. (Heineck & Faber).

References

^ Müller O. F. (1773). Vermivm terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum et testaceorum, non marinorum, succincta historia. Voluminis Imi pars Ima. pp. [1-33], 1-135. Havniae & Lipsiae. (Heineck & Faber).
^ MolluscaBase eds. (2020). MolluscaBase. Carychium O. F. Müller, 1773. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=738075 on 2020-05-01
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Ewa Stworzewicz: Miocene land snails from Belchatów (Central Poland), III: Carychiinae (Gastropoda; Pulmonata: Ellobiidae). In: Paläontologische Zeitschrift. 73(3/4), Stuttgart 1999, S. 261–276.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x Wilhelm Wenz: Gastropoda extramarina tertiaria. In: Carl Diener (Hrsg.): Fossilium catalogus, 1 Animalium. 21, Berlin 1923, S. 952–957, 1069-1420. (biodiversitylibrary.org)
^ D. C. Dourson: Four New Land Snail Species from the Southern Appalachian Mountains. In: Journal of North Carolina Academy of Science. 128(1), 2012, S. 1–10. (Abstract)
^ ^a ^b ^c Jochum, Adrienne, et al. "Three new species of Carychium OF Müller, 1773 from the Southeastern USA, Belize and Panama are described using computer tomography (CT)(Eupulmonata, Ellobioidea, Carychiidae)." ZooKeys 675 (2017): 97.(PDF)
^ Benson, W. H. "XXII.—Characters of Diplommatina, a new genus of Terrestrial Mollusks belonging to the family of Carychiadæ, and of a second species contained in it; also of a new species of Carychium inhabiting the Western Himalaya." Annals and Magazine of Natural History 4.21 (1849): 193-195. [1]
^ Hauffen, H. "Beitrage zur Grottenkunde Krain’s. Uber Carychium carinatum." Jahresheft des Vereines des krainischen Landes-Museum, Laibach 2 (1858): 96.
^ Sandberger, Fridolin. Die Land-und Süßwasser-Conchylien der Vorwelt. Text. Kreidel, 1875.
^ Benson, W. H. "XXII.—Characters of Diplommatina, a new genus of Terrestrial Mollusks belonging to the family of Carychiadæ, and of a second species contained in it; also of a new species of Carychium inhabiting the Western Himalaya." Annals and Magazine of Natural History 4.21 (1849): 193-195. [2]
^ Chitty, Edward. Descriptions of Thirty Supposed New Species and Varieties of Land and Fluviatile Shells of Jamaica: With Observations on Some Shells Already Described. RJ de Cordova, printer, 1853.
^ Von Moellendorff, O. F. "Neue Landschnecken von Java." Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29 (1897): 57-72.
^ von Möllendorff, Otto Franz. Verzeichniss der auf den Philippinen lebenden Landmollusken. 1898.
^ O. F. Müller: Vermium terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum et testaceorum, non marinorum, succincta historia. In: Volumen alterum. 1774, S. I–XXXVI, 1-214, Havniæ/Kopenhagen & Lipsiæ/Leipzig, Heineck & Faber [Online bei Biodiversity Heritage Library] (S. 125).
^ Jochum, Adrienne, et al. "Fulfilling the taxonomic consequence after DNA Barcoding: Carychium panamaense sp. n.(Eupulmonata, Ellobioidea, Carychiidae) from Panama is described using computed tomographic (CT) imaging." ZooKeys 795 (2018): 1. (PDF)
^ G. Truc: Nouveaux gastéropodes continentaux du Pliocène terminal de Celleneuve (Hérault, Sud-Est de la France). In: Documents des laboratoires de géologie de la Faculté des Sciences de Lyon. 50, 1972, S. 83–91.
^ Freyer, Custos. Ueber neu entdeckte Conchylien aus den Geschlechtern Carychium und Pterocera:(Aus dem Jännerhefte 1855 der Sitz. Ber. der math. naturw. Classe der k. Acad. der Wiss.(Bd. XV, S. 18.). 1855.
^ ^a ^b Ewa Stworzewicz, Valentin A. Prisyazhnyuk, Marcin Górka: Systematic and palaeoecological study of Miocene terrestrial gastropods from Zwierzyniec (Southern Poland). In: Annales Societatis Geologorum Poloniae. 83, 2013, S. 179–200. (PDF)

External links

Worldwide mollusc species data base
Müller, O. F. (1773-1774). Vermium terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum, et testaceorum, non marinorum, succincta historia. Vol. 1, Pars Ima: 1-136 [1773]; Pars Altera: 1-214 + index 8 pp. [1774]. Havniæ (Copenhagen) & Lipsiæ (Leipzig), Heineck & Faber
61-63. Pilsbry H. A. (October) 1894. The American species of Carychium. The Nautilus, 8(6)

[1] Müller O. F. (1773). Vermivm terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum et testaceorum, non marinorum, succincta historia. Voluminis Imi pars Ima. pp. [1-33], 1-135. Havniae & Lipsiae. (Heineck & Faber).

[WoRMS-2] MolluscaBase eds. (2020). MolluscaBase. Carychium O. F. Müller, 1773. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=738075 on 2020-05-01

[Stworzewicz-3] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Ewa Stworzewicz: Miocene land snails from Belchatów (Central Poland), III: Carychiinae (Gastropoda; Pulmonata: Ellobiidae). In: Paläontologische Zeitschrift. 73(3/4), Stuttgart 1999, S. 261–276.

[Wenz-4] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x Wilhelm Wenz: Gastropoda extramarina tertiaria. In: Carl Diener (Hrsg.): Fossilium catalogus, 1 Animalium. 21, Berlin 1923, S. 952–957, 1069-1420. (biodiversitylibrary.org)

[Dourson-5] D. C. Dourson: Four New Land Snail Species from the Southern Appalachian Mountains. In: Journal of North Carolina Academy of Science. 128(1), 2012, S. 1–10. (Abstract)

[JochumWeigand2017-6] Jochum, Adrienne, et al. "Three new species of Carychium OF Müller, 1773 from the Southeastern USA, Belize and Panama are described using computer tomography (CT)(Eupulmonata, Ellobioidea, Carychiidae)." ZooKeys 675 (2017): 97.(PDF)

[Benson1864-7] Benson, W. H. "XXII.—Characters of Diplommatina, a new genus of Terrestrial Mollusks belonging to the family of Carychiadæ, and of a second species contained in it; also of a new species of Carychium inhabiting the Western Himalaya." Annals and Magazine of Natural History 4.21 (1849): 193-195. [1]

[Haufen,_1858-8] Hauffen, H. "Beitrage zur Grottenkunde Krain’s. Uber Carychium carinatum." Jahresheft des Vereines des krainischen Landes-Museum, Laibach 2 (1858): 96.

[Sandberger1875-9] Sandberger, Fridolin. Die Land-und Süßwasser-Conchylien der Vorwelt. Text. Kreidel, 1875.

[Benson1849-10] Benson, W. H. "XXII.—Characters of Diplommatina, a new genus of Terrestrial Mollusks belonging to the family of Carychiadæ, and of a second species contained in it; also of a new species of Carychium inhabiting the Western Himalaya." Annals and Magazine of Natural History 4.21 (1849): 193-195. [2]

[Chitty1853-11] Chitty, Edward. Descriptions of Thirty Supposed New Species and Varieties of Land and Fluviatile Shells of Jamaica: With Observations on Some Shells Already Described. RJ de Cordova, printer, 1853.

[vonMöllendorff1897-12] Von Moellendorff, O. F. "Neue Landschnecken von Java." Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29 (1897): 57-72.

[vonMöllendorff1898-13] von Möllendorff, Otto Franz. Verzeichniss der auf den Philippinen lebenden Landmollusken. 1898.

[Müller1774-14] O. F. Müller: Vermium terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum et testaceorum, non marinorum, succincta historia. In: Volumen alterum. 1774, S. I–XXXVI, 1-214, Havniæ/Kopenhagen & Lipsiæ/Leipzig, Heineck & Faber [Online bei Biodiversity Heritage Library] (S. 125).

[Jochum2018-15] Jochum, Adrienne, et al. "Fulfilling the taxonomic consequence after DNA Barcoding: Carychium panamaense sp. n.(Eupulmonata, Ellobioidea, Carychiidae) from Panama is described using computed tomographic (CT) imaging." ZooKeys 795 (2018): 1. (PDF)

[Truc-16] G. Truc: Nouveaux gastéropodes continentaux du Pliocène terminal de Celleneuve (Hérault, Sud-Est de la France). In: Documents des laboratoires de géologie de la Faculté des Sciences de Lyon. 50, 1972, S. 83–91.

[Freyer1855-17] Freyer, Custos. Ueber neu entdeckte Conchylien aus den Geschlechtern Carychium und Pterocera:(Aus dem Jännerhefte 1855 der Sitz. Ber. der math. naturw. Classe der k. Acad. der Wiss.(Bd. XV, S. 18.). 1855.

[Stworzewicz2013-18] Ewa Stworzewicz, Valentin A. Prisyazhnyuk, Marcin Górka: Systematic and palaeoecological study of Miocene terrestrial gastropods from Zwierzyniec (Southern Poland). In: Annales Societatis Geologorum Poloniae. 83, 2013, S. 179–200. (PDF)

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