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{{Short description|Species of tree}}{{Use South African English|date = March 2023}}
{{Speciesbox
{{Speciesbox
| name = Quiver tree
| name = Quiver tree
| taxon = Aloidendron dichotomum
| taxon = Aloidendron dichotomum
| image = QuiverTree-Namibia-2015.JPG
| image = QuiverTree-Namibia-2015.JPG
| status =
| status = VU
| status_system =
| status_system = IUCN3.1
| status_ref = <ref name="iucn status 23 March 2023">{{cite iucn |author=[[South African National Biodiversity Institute]] |date=2022 |title=''Aloidendron dichotomum'' |volume=2022 |page=e.T140661836A140666503 |doi=10.2305/IUCN.UK.2022-1.RLTS.T140661836A140666503.en |access-date=23 March 2023}}</ref>
| status2 =
| status2_system =
| status2_ref =
| authority = ([[Francis Masson|Masson]]) Klopper & Gideon F.Sm.<ref name=WCSP_487777/>
| authority = ([[Francis Masson|Masson]]) Klopper & Gideon F.Sm.<ref name=WCSP_487777/>
| range_map2_caption = The distribution of ''Aloidendron dichotomum'' (light blue) in southern Africa
| synonyms =
{{Specieslist
| synonyms = {{Specieslist
|Aloe dichotoma|Masson
|Aloe dichotoma|Masson
|Rhipidodendrum dichotomum|(Masson) Willd.
|Rhipidodendrum dichotomum|(Masson) Willd.
Line 15: Line 20:
}}
}}
| synonyms_ref = <ref name=WCSP_487777/>
| synonyms_ref = <ref name=WCSP_487777/>
| range_map = {{maplink|frame=yes|plain=yes|from=Aloidendron dichotomum range.map}}
| range_map = The Tree Aloes of Southern Africa - Aloidendron.png
| range_map_caption = The distribution of ''Aloidendron dichotomum'' (light blue) in southern Africa
| range_map_caption = Aloidendron dichotomum range
{{leftlegend|#0094e8|Extant (resident)}}
| range_map2 = The Tree Aloes of Southern Africa - Aloidendron.png
}}
}}


'''''Aloidendron dichotomum''''', formerly '''''Aloe dichotoma''''', the '''quiver tree''' or '''kokerboom''', is a tall, branching species of [[succulent plant]], indigenous to [[Southern Africa]], specifically in the [[Northern Cape]] region of [[South Africa]], and parts of Southern [[Namibia]].
'''''Aloidendron dichotomum''''', formerly '''''Aloe dichotoma''''', the '''quiver tree''' or '''kokerboom''', is a tall, branching species of [[succulent plant]], indigenous to [[Southern Africa]], specifically in the [[Northern Cape]] province of [[South Africa]], and parts of Southern [[Namibia]].


==Naming==
==Naming==


Known as ''choje'' to the indigenous [[San people]], the quiver tree gets its English common name from the San people practice of hollowing out the tubular branches of ''Aloidendron dichotomum'' to form [[quiver]]s for their [[arrow]]s. The specific epithet ''"dichotomum"'' refers to how the stems repeatedly branch into two ("[[dichotomous branching|dichotomous]]" branching) as the plant grows.{{citation needed|date=April 2013}} This species was moved to the genus ''[[Aloidendron]]'' as ''Aloidendron dichotomum'' in 2013.<ref>https://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.76.1.2</ref>
Known as ''choje'' to the indigenous [[San people]], the quiver tree gets its English common name from the San people practice of hollowing out the tubular branches of ''Aloidendron dichotomum'' to form [[quiver]]s for their [[arrow]]s. The specific epithet ''"dichotomum"'' refers to how the stems repeatedly branch into two ("[[dichotomous branching|dichotomous]]" branching) as the plant grows.<ref name="SANBI">{{cite web |title=Aloidendron dichotomum |url=http://pza.sanbi.org/aloidendron-dichotomum |website=PlantZAfrica |publisher=South African National Biodiversity Institute |access-date=1 February 2023}}</ref> This species was moved to the genus ''[[Aloidendron]]'' as ''Aloidendron dichotomum'' in 2013.<ref name="GRACE">{{cite journal |last1=Grace |first1=O.M. |last2=Klopper |first2=R.R. |last3=Smith |first3=G.F. |title=A revised generic classification for Aloe (Xanthorrhoeaceae subfam. Asphodeloideae) |journal=Phytotaxa |date=2013 |volume=76 |issue=1 |pages=7–14 |doi=10.11646/phytotaxa.76.1.2 |s2cid=86065738 |url=https://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.76.1.2/111 |access-date=1 February 2023|hdl=2263/58378 |hdl-access=free }}</ref>


==Related species==
==Related species==
Three separate species, ''A. dichotomum'', ''[[Aloidendron pillansii|A. pillansii]]'' and ''[[Aloidendron ramosissimum|A. ramosissimum]]'' inhabit the same arid areas of the [[Richtersveld]] and the [[Namib Desert]] around the South African-Namibian border. The three have been given different ratings on the [[IUCN Red List|IUCN Red List of Threatened Species]]: 'vulnerable' for ''A. dichotomum'', 'critically endangered' for ''A. pillansii'' and 'endangered' for ''A. ramossisimum''.
Three separate species, ''A. dichotomum'', ''[[Aloidendron pillansii|A. pillansii]]'' and ''[[Aloidendron ramosissimum|A. ramosissimum]]'' inhabit the same arid areas of the [[Richtersveld]] and the [[Namib Desert]] around the South African-Namibian border. The three have been given different ratings on the [[IUCN Red List|IUCN Red List of Threatened Species]]: 'vulnerable' for ''A. dichotomum'', 'critically endangered' for ''A. pillansii'' and 'endangered' for ''A. ramossisimum''.{{cn|date=March 2023}}


The three species can be distinguished. In ''A. pillansii'', the [[inflorescence]]s hang from below the lowest leaves, rather than growing erect. ''A. ramosissimum'' is considerably smaller - rarely reaching more than 2&nbsp;m in height - and assumes a more shrub-like shape. While there is a gradation between tree-like ''A. dichotomum'' and the shrubby ''A. ramosissimum'', the relatively unique ''A. pillansii'' population is separated by a different flowering time and therefore does not interbreed with the other two species.<ref name=Court2010>{{Cite book |last=Court |first=D. |year=2010 |title=Succulent Flora of Southern Africa |location=Cape Town |publisher=Struik Nature |isbn=978-1-77007-587-0 }}</ref>
The three species can be distinguished as follows: In ''A. pillansii'', the [[inflorescence]]s hang from below the lowest leaves, rather than growing erect. ''A. ramosissimum'' is considerably smaller—rarely reaching more than 2&nbsp;m in height—and assumes a more shrub-like shape. While there is a gradation between tree-like ''A. dichotomum'' and the shrubby ''A. ramosissimum'', the relatively unique ''A. pillansii'' population is separated by a different flowering time and therefore does not interbreed with the other two species.<ref name=Court2010>{{Cite book |last=Court |first=D. |year=2010 |title=Succulent Flora of Southern Africa |location=Cape Town |publisher=Struik Nature |isbn=978-1-77007-587-0 }}</ref>


==Distribution and conservation==
==Distribution and conservation==
One of the few examples of spontaneous forests of ''A. dichotomum'' is the [[Quiver Tree Forest]], about 14&nbsp;km north of [[Keetmanshoop]], in [[Namibia]]. Another is located in the Northern Cape of South Africa at Gannabos.
One of the few examples of spontaneous forests of ''A. dichotomum'' is the [[Quiver Tree Forest]], about 14&nbsp;km north of [[Keetmanshoop]], in [[Namibia]]. Another is located in the Northern Cape of South Africa at Gannabos.{{cn|date=March 2023}}


Throughout much of its range this species is in decline. Modeling of ''Aloidendron dichotomum'' in [[South Africa]] and [[Namibia]] has contributed to understanding the needs of protected areas in response to climate change. Modelled range declines in this species due to climate change have recently been confirmed by field surveys.<ref>{{Cite journal |last1=Foden |first1=Wendy |author-link=Wendy Foden |last2=Midgley |first2=Guy F. |last3=Hughes |first3=Greg |last4=Bond |first4=William J. |last5=Thuiller |first5=Wilfried |last6=Hoffman |first6=M. Timm |last7=Kaleme |first7=Prince |last8=Underhill |first8=Les G. |last9=Rebelo |first9=Anthony |display-authors=8|year=2007 |title=A changing climate is eroding the geographical range of the Namib Desert tree ''Aloe'' through population declines and dispersal lags |journal=Diversity and Distributions |volume=13 |issue=5 |pages=645–653 |url=http://www4.nau.edu/direnet/publications/publications_f/files/Foden-2007.pdf |accessdate=16 July 2011 |name-list-style=amp |doi=10.1111/j.1472-4642.2007.00391.x}}</ref>
Throughout much of its range this species is in decline. Modelling of ''Aloidendron dichotomum'' in [[South Africa]] and [[Namibia]] has contributed to understanding the needs of protected areas in response to climate change. Modelled range declines in this species due to climate change have recently been confirmed by field surveys.<ref>{{Cite journal |last1=Foden |first1=Wendy |author-link=Wendy Foden |last2=Midgley |first2=Guy F. |last3=Hughes |first3=Greg |last4=Bond |first4=William J. |last5=Thuiller |first5=Wilfried |last6=Hoffman |first6=M. Timm |last7=Kaleme |first7=Prince |last8=Underhill |first8=Les G. |last9=Rebelo |first9=Anthony |display-authors=8|year=2007 |title=A changing climate is eroding the geographical range of the Namib Desert tree ''Aloe'' through population declines and dispersal lags |journal=Diversity and Distributions |volume=13 |issue=5 |pages=645–653 |url=http://www4.nau.edu/direnet/publications/publications_f/files/Foden-2007.pdf |accessdate=16 July 2011 |name-list-style=amp |doi=10.1111/j.1472-4642.2007.00391.x|s2cid=53662810 }}</ref>


==Cultivation==
==Cultivation==
''Aloidendron dichotomum'' is cultivated in arid areas around the world, for use in landscaping. The slow growth rate and relative rarity of the plant make it a particularly expensive specimen. It is also relatively difficult to keep outside of its natural habitat.
''Aloidendron dichotomum'' is cultivated in arid areas around the world, for use in landscaping. The slow growth rate and relative rarity of the plant make it a particularly expensive specimen. It is also relatively difficult to keep outside of its natural habitat.{{cn|date=March 2023}}


In cultivation it requires extremely well-drained coarse mineral sand (preferably with some loam and bone meal to keep it active and growing), full sun, good aeration and extremely little water - primarily in the winter (as it mainly occurs in winter rainfall desert). In the (rare) event that it is under-watered, the leaves will curl up and die off at the tips; this is not fatal, but indicates that it is relatively dry.
In cultivation it requires extremely well-drained coarse mineral sand (preferably with some loam and bone meal to keep it active and growing), full sun, good aeration and extremely little water - primarily in the winter (as it mainly occurs in winter rainfall desert areas). In the (rare) event that it is under-watered, the leaves will curl up and die off at the tips; this is not fatal, but indicates that it is relatively dry.{{cn|date=March 2023}}


It is unusually prone to aphids and insect infections in between its leaves, and this is exacerbated whenever there is not full sun and constant fresh air movement. Indoor plants require frequent treatment for these pests. Fungicide can also be added occasionally, to protect from rot.
It is unusually prone to aphids and insect infections in between its leaves, and this is exacerbated whenever there is not full sun and constant fresh air movement. Indoor plants require frequent treatment for these pests. Fungicide can be added occasionally, to protect the plant from rot.{{cn|date=March 2023}}


It can be propagated from seed and (with more difficulty) from cuttings or truncheons. Cuttings need to be thoroughly dried for several weeks in a shaded area, before being planted.<ref>{{cite web|url=http://www.plantzafrica.com/plantab/aloedichotoma.htm|title=Aloe dichotoma|publisher=|accessdate=26 September 2016}}</ref>
It can be propagated from seed and (with more difficulty) from cuttings or truncheons. Cuttings need to be thoroughly dried for several weeks in a shaded area before being planted.<ref name="SANBI"/>


==In popular culture==
==Movie References==
Miachel Benson's book "Space Odyssey: Stanley Kubrick, Arthur C. Clarke, and the Making of a Masterpiece" (2018) briefly discusses how the kokerboom piqued the interest of film director, Stanley Kubrick. The book claims that several protected kokerboom trees were cut down so they could be used to film "2001: A Space Odyssey".<ref>Space Odyssey: Stanley Kubrick, Arthur C. Clarke, and the Making of a Masterpiece. (2018) p. 277. http://www.simonandschuster.com/books/Space-Odyssey/Michael-Benson/9781501163937</ref>
Michael Benson's book ''Space Odyssey: Stanley Kubrick, Arthur C. Clarke, and the Making of a Masterpiece'' (2018) briefly discusses how the kokerboom piqued the interest of film director, [[Stanley Kubrick]]. The book claims that several protected kokerboom trees were cut down so they could be used to film ''[[2001: A Space Odyssey (film)|2001: A Space Odyssey]]''.<ref>{{Cite book |last=Benson |first=Michael |url=https://www.simonandschuster.com/books/Space-Odyssey/Michael-Benson/9781501163944 |title=Space Odyssey: Stanley Kubrick, Arthur C. Clarke, and the Making of a Masterpiece |date=2019-04-23 |publisher=[[Simon & Schuster]] |isbn=978-1-5011-6394-4 |pages=277 |language=en}}</ref>


==Gallery==
==Gallery==
<gallery mode=packed heights=90px>
<gallery mode="packed" heights="120px">
Image:Áloe-aljaba (Aloe dichotoma), parque nacional de Namib-Naukluft, Namibia, 2018-08-05, DD 39.jpg|Quiver tree in the [[Namib-Naukluft National Park]], Namibia.
Image:Áloe-aljaba (Aloe dichotoma), parque nacional de Namib-Naukluft, Namibia, 2018-08-05, DD 39.jpg|Quiver tree in the [[Namib-Naukluft National Park]], Namibia.
Image:Aloe dichotoma in Phoenix.jpg|In bloom.
Image:Aloe dichotoma in Phoenix.jpg|In bloom.
Line 54: Line 61:
Image:Aloe-dichotoma-young-time-comparison.jpg|Growth of a young plant from May to August.
Image:Aloe-dichotoma-young-time-comparison.jpg|Growth of a young plant from May to August.
Image:Koecherbaum Fish River Canyon anagoria.JPG|Quiver tree at Fish River Canyon, Namibia.
Image:Koecherbaum Fish River Canyon anagoria.JPG|Quiver tree at Fish River Canyon, Namibia.
<br>
Image:Köcherbaumwald-01.jpg|Quiver Tree Forest near Keetmanshoop, Namibia, in the evening.
Image:Köcherbaumwald-01.jpg|Quiver Tree Forest near Keetmanshoop, Namibia, in the evening.
Image:Kokerboom.jpg|[[Sociable Weaver]] (''Philetairus socius'') nest in a Quiver tree. Northern Cape, South Africa.
Image:Kokerboom.jpg|[[Sociable Weaver]] (''Philetairus socius'') nest in a Quiver tree. Northern Cape, South Africa.
Line 71: Line 79:


==External links==
==External links==
{{Commons category|Aloe dichotoma}}
* [http://www.desert-tropicals.com/Plants/Asphodelaceae/Aloe_dichotoma.html Desert-tropicals.com profile]
* [http://www.desert-tropicals.com/Plants/Asphodelaceae/Aloe_dichotoma.html Desert-tropicals.com profile] {{Webarchive|url=https://web.archive.org/web/20170510232251/http://www.desert-tropicals.com/Plants/Asphodelaceae/Aloe_dichotoma.html |date=2017-05-10 }}
* [http://www.plantzafrica.com/plantab/aloedichotoma.htm PlantZAfrica.com profile]
* [http://www.plantzafrica.com/plantab/aloedichotoma.htm PlantZAfrica.com profile]
* {{AfricanPlants|Aloidendron dichotomum}}
* {{AfricanPlants|Aloidendron dichotomum}}
{{Commons category|Aloe dichotoma|<br/>Aloe dichotoma|position=left}}


{{Taxonbar|from1=Q42729505|from2=Q161263}}
{{Taxonbar|from1=Q42729505|from2=Q161263}}

Latest revision as of 12:53, 12 May 2024

Quiver tree
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Order: Asparagales
Family: Asphodelaceae
Subfamily: Asphodeloideae
Genus: Aloidendron
Species:
A. dichotomum
Binomial name
Aloidendron dichotomum
(Masson) Klopper & Gideon F.Sm.[2]
Map
Aloidendron dichotomum range
  Extant (resident)
The distribution of Aloidendron dichotomum (light blue) in southern Africa
Synonyms[2]
  • Aloe dichotoma Masson
  • Rhipidodendrum dichotomum (Masson) Willd.
  • Aloe ramosa Haw.
  • Aloe montana Schinz
  • Aloe dichotoma var. montana (Schinz) A.Berger

Aloidendron dichotomum, formerly Aloe dichotoma, the quiver tree or kokerboom, is a tall, branching species of succulent plant, indigenous to Southern Africa, specifically in the Northern Cape province of South Africa, and parts of Southern Namibia.

Naming

[edit]

Known as choje to the indigenous San people, the quiver tree gets its English common name from the San people practice of hollowing out the tubular branches of Aloidendron dichotomum to form quivers for their arrows. The specific epithet "dichotomum" refers to how the stems repeatedly branch into two ("dichotomous" branching) as the plant grows.[3] This species was moved to the genus Aloidendron as Aloidendron dichotomum in 2013.[4]

[edit]

Three separate species, A. dichotomum, A. pillansii and A. ramosissimum inhabit the same arid areas of the Richtersveld and the Namib Desert around the South African-Namibian border. The three have been given different ratings on the IUCN Red List of Threatened Species: 'vulnerable' for A. dichotomum, 'critically endangered' for A. pillansii and 'endangered' for A. ramossisimum.[citation needed]

The three species can be distinguished as follows: In A. pillansii, the inflorescences hang from below the lowest leaves, rather than growing erect. A. ramosissimum is considerably smaller—rarely reaching more than 2 m in height—and assumes a more shrub-like shape. While there is a gradation between tree-like A. dichotomum and the shrubby A. ramosissimum, the relatively unique A. pillansii population is separated by a different flowering time and therefore does not interbreed with the other two species.[5]

Distribution and conservation

[edit]

One of the few examples of spontaneous forests of A. dichotomum is the Quiver Tree Forest, about 14 km north of Keetmanshoop, in Namibia. Another is located in the Northern Cape of South Africa at Gannabos.[citation needed]

Throughout much of its range this species is in decline. Modelling of Aloidendron dichotomum in South Africa and Namibia has contributed to understanding the needs of protected areas in response to climate change. Modelled range declines in this species due to climate change have recently been confirmed by field surveys.[6]

Cultivation

[edit]

Aloidendron dichotomum is cultivated in arid areas around the world, for use in landscaping. The slow growth rate and relative rarity of the plant make it a particularly expensive specimen. It is also relatively difficult to keep outside of its natural habitat.[citation needed]

In cultivation it requires extremely well-drained coarse mineral sand (preferably with some loam and bone meal to keep it active and growing), full sun, good aeration and extremely little water - primarily in the winter (as it mainly occurs in winter rainfall desert areas). In the (rare) event that it is under-watered, the leaves will curl up and die off at the tips; this is not fatal, but indicates that it is relatively dry.[citation needed]

It is unusually prone to aphids and insect infections in between its leaves, and this is exacerbated whenever there is not full sun and constant fresh air movement. Indoor plants require frequent treatment for these pests. Fungicide can be added occasionally, to protect the plant from rot.[citation needed]

It can be propagated from seed and (with more difficulty) from cuttings or truncheons. Cuttings need to be thoroughly dried for several weeks in a shaded area before being planted.[3]

[edit]

Michael Benson's book Space Odyssey: Stanley Kubrick, Arthur C. Clarke, and the Making of a Masterpiece (2018) briefly discusses how the kokerboom piqued the interest of film director, Stanley Kubrick. The book claims that several protected kokerboom trees were cut down so they could be used to film 2001: A Space Odyssey.[7]

[edit]

See also

[edit]

References

[edit]
  1. ^ South African National Biodiversity Institute (2022). "Aloidendron dichotomum". IUCN Red List of Threatened Species. 2022: e.T140661836A140666503. doi:10.2305/IUCN.UK.2022-1.RLTS.T140661836A140666503.en. Retrieved 23 March 2023.
  2. ^ a b "Aloidendron dichotomum". World Checklist of Selected Plant Families. Royal Botanic Gardens, Kew. Retrieved 16 October 2017.
  3. ^ a b "Aloidendron dichotomum". PlantZAfrica. South African National Biodiversity Institute. Retrieved 1 February 2023.
  4. ^ Grace, O.M.; Klopper, R.R.; Smith, G.F. (2013). "A revised generic classification for Aloe (Xanthorrhoeaceae subfam. Asphodeloideae)". Phytotaxa. 76 (1): 7–14. doi:10.11646/phytotaxa.76.1.2. hdl:2263/58378. S2CID 86065738. Retrieved 1 February 2023.
  5. ^ Court, D. (2010). Succulent Flora of Southern Africa. Cape Town: Struik Nature. ISBN 978-1-77007-587-0.
  6. ^ Foden, Wendy; Midgley, Guy F.; Hughes, Greg; Bond, William J.; Thuiller, Wilfried; Hoffman, M. Timm; Kaleme, Prince; Underhill, Les G.; et al. (2007). "A changing climate is eroding the geographical range of the Namib Desert tree Aloe through population declines and dispersal lags" (PDF). Diversity and Distributions. 13 (5): 645–653. doi:10.1111/j.1472-4642.2007.00391.x. S2CID 53662810. Retrieved 16 July 2011.
  7. ^ Benson, Michael (2019-04-23). Space Odyssey: Stanley Kubrick, Arthur C. Clarke, and the Making of a Masterpiece. Simon & Schuster. p. 277. ISBN 978-1-5011-6394-4.
[edit]